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Workshops

Introduction Upcoming Activities Past Activities


Introduction


To advance specific areas of speciation research, the programme will organize seven workshops on selected topics of contemporary interest. To facilitate bridge-building between disparate approaches to speciation research, these workshops will bring together participants of different backgrounds for a few days of intense discussions and presentations. The seven workshops are described in further detail below, including contact information of the principal organizers.



Upcoming Activities


Title: Hybridisation and speciation
Date: 23-26 October 2011
Location: Gregynog Hall, Wales, UK
Main organizer: Roger Butlin (r.k.butlin@sheffield.ac.uk)
Other organizers: Mike Ritchie, Jacek Szymura, Åke Brännström, and Ulf Dieckmann


Expected number of participants: 10 invited speakers; 40 participants


Application procedure: Prospective partcipants should send a brief CV with a list of publications (if any) together with a one-paragraph statement of motivation. Applications will be accepted between eight and two months before the event. Later applications can be considered if places are available.


Program outline: The first 4 half-day sessions will begin with a keynote presentation and be followed by discussions structured around the pre-submitted statements and questions. All participants will be expected to come prepared to expand upon and defend their statements/questions and challenge others. The final half-day session will attempt to reach agreement on statements of established principles and priorities for future work (and so will outline the proposed publication).


Summary: Hybridisation, mating and offspring production involving genetically differentiated populations, and introgression, the passage of genes from one genetically distinct population into another, are key processes in many models of speciation. They may occur in defined regions following secondary contact or as a result of local adaptation, in hybrid zones, or more broadly across species distributions. There may be a stable balance between hybridisation and selection or hybridisation may initiate the collapse of differentiation or the origin of new species. Differential introgression offers crucial insights into the role of selection during the evolution of reproductive isolation. With a sophisticated theoretical foundation developed in the last 30 years, recent technological advances now offer particularly exciting opportunities to study the consequences of hybridisation and so to advance understanding of the process of speciation. The workshop will include a small number of keynote lectures but most of the time will be devoted to small-group discussions. To focus these discussions, we will ask participants to send, in advance of the meeting, statements of key principles or empirical patterns which they consider to be firmly established and key questions that remain to be answered, theoretically or empirically. These will be collated and used to structure the meeting. They will also form the basis for a publication after the meeting.



Title: Community evolution and speciation
Date: 2012
Location: Umeå University, Umeå, Sweden
Main organizers: Åke Brännström (ake.brannstrom@math.umu.se), Ulf Dieckmann (dieckmann@iiasa.ac.at)


Expected number of participants: 10 invited speakers; open number of participants at own expense


Application procedure: Prospective partcipants should send a brief CV with a list of publications (if any) together with a one-paragraph statement of motivation. Applications will be accepted between eight and two months before the event. Later applications can be considered if places are available.


Program outline: The proposed workshop will include talks by leading figures in the field. These will be integrated with discussion sessions.


Summary: The traditional focus of speciation research has been on the processes that can create two species out of one. While these processes underpin the creation of biological diversity, it is important to recognize that they only meant to operate under specific, relatively simple ecological conditions. These conditions can be differences in the external environments to which the two species adapt, but in many cases the niches to which species are adapting exist only in the context of a community or ecosystem embedding. Ultimately, all ecological niches are constructed, rather than preordained. Understanding how community evolution endogenously creates new niches for other species to fill is therefore an important problem. Interest in this question has mounted during recent years. In this workshop we will bring together scientists with backgrounds in classical speciation research and community ecology for discussions of a common theme, thus aiming to bridge between these disparate approaches to understanding the complexity of ecological communities.



Title: Genetics and genomics of speciation
Date: Summer 2012 or 2013
Location: ETHZ Conference Centre Stefano Franscini (CSF) on Monte Verita above Ascona (Ticino)
Main organizer: Ole Seehausen (ole.seehausen@eawag.ch)
Other organizers: Radka Storchova (Frospects steering-committee member), Roger Butlin (Frospects steering-committee member), Elena Conti (Uni Zurich), Laurent Excoffier (Uni Bern), Walter Salzburger (Uni Basel), Alex Widmer (ETH Zurich), Marta Barluenga (Eawag), Irene Keller (Eawag), Arjun Sivasundar (Uni Bern), Piet Spaak (Eawag), Barbara Taborsky (Uni Bern), Ulf Dieckmann (Frospects chair), Åke Brännström (Frospects co-chair)


Expected number of participants: 10-15 invited speakers; upper limit of 50 or 70 participants, depending on venue


Application procedure: Prospective partcipants should send a brief CV with a list of publications (if any) together with a one-paragraph statement of motivation. Applications will be accepted between eight and two months before the event. Later applications can be considered if places are available.


Program outline: We wish to review recent advances in the genetics and genomics of speciation, and discuss how these can be integrated with advances in ecology to test theoretical models of speciation, and to inform further.


Summary: The recent renaissance in speciation research was driven significantly by advances in molecular phylogenetics and population genetics, and the integration of these disciplines with evolutionary ecology, community ecology and biogeography. A growing data analysis machinery permits increasingly detailed reconstruction of gene flow and selection. The technological advances of the genomics era are beginning to open windows into genome-wide effects of divergent adaptation, and the functional basis of species divergence and incompatibility. Finally, whole genomes of some speciation model organisms are being sequenced and annotated. Speciation researchers are beginning to capitalize on these developments. However, most genomic species differences arise after speciation, and to understand the process, speciation has to be studied in the making. The impact of new technologies on the field is therefore strictly dependent on the quality of natural history and ecology. The integration of genomics with ecology is perhaps the most important challenge to speciation research in the next decade. In this workshop we want to review recent advances in, and identify future needs for the integration of these disciplines in speciation research. We hope to identify ways by which their integration can advance our abilities both for testing theoretical models, and for informing further theory development.



Title: Modes of diversification
Date: 2013
Location: To be announced
Main organizers: Åke Brännström (ake.brannstrom@math.umu.se), Ulf Dieckmann (dieckmann@iiasa.ac.at)


Expected number of participants: 10-15 invited speakers; upper limit of 40 participants


Application procedure: Prospective participants should send a brief CV with a list of publications (if any) together with a one-paragraph statement of motivation. Applications will be accepted between eight and two months before the event. Later applications can be considered if places are available.


Program outline: The proposed workshop will include talks by leading figures in the field. These will be integrated with discussion sessions.


Summary: The traditional ‘standard model’ of speciation rests on the assumption of geographic isolation. The trigger for speciation in this standard model is geographic isolation. It is for this reason that the distinction between allopatric speciation and sympatric speciation has taken centre stage in the speciation debate. Strictly speaking, the terms allopatric and sympatric refer only to the biogeographic patterns in the distribution of a species. By disentangling pattern and process, several additional modes of speciation can be identified. These include ecological speciation, competitive speciation, and adaptive speciation, which are all closely related. The aim of this workshop is to discuss different modes of speciation from theoretical and empirical angles. Particular focus will be given to signatures of speciation, i.e., to methods for inferring the speciation mode based on information on ecological, genetic, and spatial diversification.
 


Title: Behaviour and speciation
Date: 2013
Location: Centre for Ecological and Evolutionary Synthesis (CEES), University of Oslo
Organizers: Glenn-Peter Sætre (g.p.satre@bio.uio.no)
Other organizers: Tentatively: Nils Chr Stenseth (Univ Oslo), Stein Are Sæther (Univ Oslo), Øistein Haugsten Holen (Univ Oslo), Asbjørn Vøllestad (Univ Oslo), Åke Brännström, Ulf Dieckmann


Expected number of participants: 10 invited speakers; 40 participants incl. students


Application procedure: Prospective partcipants should send a brief CV with a list of publications (if any) together with a one-paragraph statement of motivation. Applications will be accepted between eight and two months before the event. Later applications can be considered if places are available.


Program outline: Tentatively, lectures focusing on key questions on behavioural aspects in speciation research.
 
Summary: Many populations are thought to be reproductively isolated from other such populations (i.e. they are different species) by choice. That is, individuals from the differentiated populations are capable of mating with each other and produce fertile offspring but they do not do so due to a lack of sexual attraction; the populations are reproductively isolated by pre-mating barriers. This is but one example of the potential importance of behavioural mechanisms in hindering gene flow and hence in speciation. In this workshop we intend to explore behavioural interactions between differentiated populations in secondary contact and sympatry and address their importance in population divergence, convergence and speciation, focusing on both empirical and theoretical approaches. Relevant topics include: learning, genes and species recognition; the behavioural ecology of reinforcement; intra- and intersexual signalling and communication in hybrid zones.



Past Activities


Title: Niche theory and speciation
Date: 29-31 August 2011
Location: Lake Balaton, Hungary
Main organizer: Géza Meszéna (geza.meszena@elte.hu)
Other organizers: Liz Pásztor, András Szilágyi, Ake Brännström, and Ulf Dieckmann


Number of participants: 20 invited speakers; 20 participants


Program outline: The program was developed interactively with the invited lecturers considering the abstract below as a point of departure. The speakers were approached to establish their favorite subject to lecture on as well as to hear about.


Summary: The workshop's aim was to integrate theories of generation and maintenance of species diversity. The Darwinian idea for the origin of species connects divergence to weakening struggle for existence. It is inherently related to Darwin’s view on reduced competition between species with different roles in the economy of nature. In modern terms, Darwinian speciation is necessarily based on niche segregation. In particular, adaptation to different conditions (“habitats”) and to different resources can be seen as two complementary ways of reducing competition. This way, the different modes of speciation can be seen as different realizations of the same underlying phenomenon. Over the last decade robust empirical evidence has been accumulating on the mechanisms maintaining species coexistence and on frequency-dependent selection, ecological divergence as well as on sympatric speciation. We have invited people working on both the empirical and theoretical aspects of niche segregation, phenotypic evolution and genetics of speciation to develop connections between the ecological and genetic theories of speciation.



Title: Evolution of divergence and speciation models of specific systems
Date: 3-6 August 2010
Location: Hólar University College, Hólar, Iceland
Main organizers: Bjarni K. Kristjánsson (bjakk@holar.is), Skúli Skúlason (skuli@holar.is)
Other organizers: Sigurður S. Snorrason, Åke Brännström, and Ulf Dieckmann


Number of participants: 14 invited speakers; 40 participants


Program outline: Program consisted of 30 min. presentations by invited speakers followed by ample time for discussion. Other participants were offered to present posters and to make short verbal presentations in special sessions. Special discussion periods were organized every day in a relaxed environment to ensure creative progress and output. An effort was made to connect individuals well and link across fields and study systems.


Summary: Historically, ideas and models of speciation have focused on either allopatric or sympatric divergence. Recent advances in theoretical, ecological, molecular and developmental approaches have opened new and more detailed avenues for the understanding speciation processes. Systems where intra-specific morphs and populations exist are particularly useful for the study of speciation. Speciation models and empirical studies of specific systems were examined and compared. Firstly, the function of ecology was examined for phenotypic and reproductive divergence, along with reproductive styles and mating biology. Secondly, it was explored how genetic and environmental (plastic) mechanisms work and interact during divergence, e.g. at early stages of divergence and in the evolution of reproductive isolation. Thirdly, theoretical models that have been constructed in relation to specific systems were examined. The workshop has focused on specific verbal and theoretical models and empirical examples, reviewing recent advances in this field and comparing different systems of organisms to explore if new and common theoretical and empirical patterns are being realized.
 

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Last updated: 06 Sep 2011